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In Drosophila Melanogaster, when and how was the carnation (car) gene discovered?

In Drosophila Melanogaster, when and how was the carnation (car) gene discovered?

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VPS33B protein is a homologue of yeast class C vacuolar protein sorting protein Vps33p that is involved in the biogenesis function of vackules. Mutation in Vps33p homologue result in different phenotype for example mutation Drosophila melanogaster car gene causes the carnation eye colour mutant. Thus in Drosophil, a hypomorphic allele of the Vps33 homologue carnation causes the carnation eye colour phenotype. The car gene product localizez to endosomel compartments and is a homolog of SM regulators of membrane fusion.

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