Step-by-Step Solution
The un-3 locus in Neurospora is near the centromere on chromosome 1. Crossovers between the un-3 locus and centromere are quite rare. The ad-3 locus is on the other side of the centromere and on the same chromosome. Crossovers between ad-3 and the centromere occur in about 20% of meioses.
a) Single crossovers between a gene and its centromere lead to tetratypes, which indicates the occurrence of second division segregation. Hence, a total of 20% of asci should show second division segregation. The remaining 80% of asci will show first division segregation. The representative asci can be as follows:
For all the different asci given, the upside down order will also occur.
b) When a un-3 ad-3 strain was crossed with a wild type, the results were:
The aborted spores could result from a crossing over event within an inversion of the wild type. When crossing over occurs within heterozygous inversions, it leads to unbalanced chromosomes and nonviable spores.
This idea can be tested by using the wild type from Hawaii in mapping experiments. Other markers on chromosome 1 can be crossed with the standard strain and observed for any alterations in map distances.
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